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Enabling replication between AD and Exchange 5.5 directory

We currently have an Exchange 5.5 Org with four sites, and we're about to start upgrading to Win2k and Exchange 2000. Is one ADC needed per 5.5 site, or is just one needed for the entire 5.5 organization?
To enable replication between Active Directory and your Exchange 5.5 directory, each mixed site in your organization must have a two-way Connection Agreement (CA) configured. The server specified under Exchange Server information on the Connections tab of the CA properties must be either an Exchange 5.5 computer or an SRS server in the mixed site.

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